Queen Beryl

Serving as the primary antagonist in the early stages of the Sailor Moon saga, this powerful leader commands both the Dark Kingdom and the four formidable Shitenou. While the anime depicts her as the ruler of the Negaverse, her manga counterpart possesses a much more tragic backstory involving the invasion of the Silver Millennium. Ultimately, her reign ends when Sailor Moon defeats her, resulting in her banishment back to her dark dimension.